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Abstract

This study is the first comparison of the morphology of pollen grains in ten cultivars of three species of the Taxus,

Torreya nucifera and Cephalotaxus harringtonia var. drupacea genera. The material came from the Botanical

Garden of Adam Mickiewicz University in Poznań, Poland. Each measurement sample consisted of 50 pollen

grains. In total, 750 pollen grains were analyzed. Light and electron scanning microscopy was used for the morphometric

observation and analysis of pollen grains. The pollen grains were inaperturate and classified as small

and medium-sized. They were prolate-spheroidal, subprolate to prolate in shape. The surface of the exine was

microverrucate-orbiculate, perforate in Cephalotaxus harringtonia var. drupacea, granulate-orbiculate, perforate

in all Taxus taxa and granulate-microverrucate-orbiculate, perforate in Torreya. The orbicules were rounded to

oval in surface view, and the size was considerably diversified. The pollen features were insufficient to distinguish

between individual Taxus members – only groups were identified. The values of the coefficient of variability of

three features (LA, SA and LA/SA) were significantly lower than the orbicule diameter. The pollen surface of all

Taxus specimens was similar, so it was not a good identification criterion. The pollen grains of the Taxus taxa

were smaller and had more orbicules than Cephalotaxus and Torreya. Palynological studies provided taxonomic

support for recognition of two different genera of the Cephalotaxaceae and Taxaceae families, which are closely

related.

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Authors and Affiliations

Joanna Bykowska
Małgorzata Klimko
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Abstract

In this autobiographical note I describe my childhood and early University days in London, including the initiation of research on the Cenomanian chalks of southern England under the supervision of the late Jake Hancock, who was to become the closest of friends and collaborators for nearly 40 years. Appointment to a teaching post in Oxford in 1967 led, eventually, to the directorship of the Oxford University Museum of Natural History in 2003, until retirement in 2010. It was my good fortune to travel widely in connection with research on the Cretaceous across Europe and the United States, but particularly in KwaZulu Natal in South Africa, leading to a career long collaboration with Herbert Klinger (Cape Town). Collaboration has been the key to my research, collaboration with Jake and Herbie, and many others, including Bill Cobban, Andy Gale, Pierre Juignet, Herbert Summesberger, Irek Walaszczyk, and Willy Wright. These collaborations led to publications that dealt with ammonite faunas from The Antarctic Peninsula to Greenland, and from the United States Western Interior to Australia, as listed below.
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Authors and Affiliations

William James Kennedy
1 2

  1. Oxford University Museum of Natural History, Parks Road, Oxford OX1 3PW, United Kingdom
  2. Department of Earth Sciences, South Parks Road, Oxford OX1 3AN, United Kingdom
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Abstract

New information about presence and features of some Lecanora species as well as their ecology and distribution in Antarctica are provided. Lecanora dispersa (Pers.) Sommerf. is confirmed to occur in the Antarctic region; L. sverdrupiana Řvst. is recorded for the first time from maritime Antarctica; L. torrida Vain. is reported as new for that Antarctic area and for the southern hemisphere. An attempt to summarize the present state of knowledge for the genus Lecanora in the Antarctic region is made. Several species, which require more in depth studies, are briefly discussed and an up-to-date list of species occurring in Antarctica is included.

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Authors and Affiliations

Lucyna Śliwa
Maria Olech
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Abstract

Five species of Tardigrada were found in two moss samples collected from the Hornsund area (Ariekammen, Spitsbergen) including one new to science. The new species, Isohypsibius karenae sp. n., differs from the other similar congeners mainly by having a different type of cuticular sculpture, a different macroplacoid length sequence, by the presence of lunules and cuticular bars under claws as well as by some morphometric characters. The current study increases the number of Isohypsibius species known from Svalbard to thirteen.
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Authors and Affiliations

Krzysztof Zawierucha
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Abstract

The first records from samples from the IceAGE cruise ME85/3 in 2011 include seven species of Caudofoveata with a distribution range in Icelandic waters. From this first cruise of the project, two new records for Iceland have been registered. Psilodens balduri sp. n. is new to science and Falcidens halanychi , with a known distribution in the American North−Atlantic, is new to Iceland. The current study thus increases the number of known caudofoveate species around Iceland to nine.
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Authors and Affiliations

Nina T. Mikkelsen
Christiane Todt
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Abstract

During August–September 2011, scientists aboard the R/V Meteor sampled marine animals around Iceland for the IceAGE project (Icelandic marine Animals: Genetics and Ecology). The last sample was taken at a site known as “The Rose Garden” off north− eastern Iceland and yielded a large number of two species of Proneomenia (Mollusca, Aplacophora, Solenogastres, Cavibelonia, Proneomeniidae). We examined isolated sclerites, radulae, and histological section series for both species. The first, Proneomenia sluiteri Hubrecht, 1880, was originally described from the Barents Sea. This is the first record of this species in Icelandic waters. However, examination of aplacophoran lots collected during the earlier BIOICE campaign revealed additional Icelandic localities from which this species was collected previously. The second represents a new species of Proneomenia, which, unlike other known representatives of the genus, broods juveniles in the mantle cavity. We provide a formal description, proposing the name Proneomenia custodiens sp. n. Interestingly, the sclerites of brooded juveniles are scales like those found in the putatively plesiomorphic order Pholidoskepia rather than hollow needles like those of the adults of this species. Cytochrome c oxidase subunit I (COI) DNA barcode sequences are provided for both species of Proneomenia .
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Authors and Affiliations

Christiane Todt
Kevin M. Kocot
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Abstract

This paper presents new records of stenothoids from the Scotia Arc (West Antarctic). Altogether twenty species were recorded, two of which are reported in the West Antarctic for the first time. In addition, two species are here recorded for the first time since their description. New data on distribution are supplemented by taxonomical remarks on the collected species.
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Authors and Affiliations

Anna Jażdżewska
Traudl Krapp-Schickel
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Abstract

Starting from a subjective viewpoint on the decreasing interest in invertebrate fossil taxonomy, this essay discusses its importance in palaeobiological studies exemplified with cases from the palaeobiogeography and palaeoecology of rugose corals, and aims at provoking a discussion on the topic. The possible causes of this negative declining trend include inherent problems of palaeontological taxonomy, and changing systems in science and higher education.
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Authors and Affiliations

Jerzy Fedorowski
1

  1. Institute of Geology, Adam Mickiewicz University, Bogumiła Krygowskiego 12, 61-680 Poznań, Poland
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Abstract

The achene morphology and pericarp anatomy of 12 taxa representing three genera ( Anemone, Hepatica, and Pulsatilla) of the subtribe Anemoninae were investigated using microtome and light microscopy to evaluate the taxonomic implications of achene characters. The achenes of Anemone were elliptical or obovoid and beaked, whereas the achene of Hepatica and Pulsatilla were obovoid and elliptical, respectively. Noticeable variations in both quantitative and qualitative features of achenes were observed among the species of the three genera. One-way analysis of variance indicated that the quantitative achene variables among the species were highly significant (P<0.001). Pearson’s correlation coefficient also showed a significant correlation between different achene variables. The pericarp structure, particularly the number of cell layers and cell forms in the exocarp and endocarp, seems to be very useful for species delimitation in Anemone and Hepatica. The nature of the endotesta could provide substantial proof for sub-generic classification in Anemone. Unweighted paired group analysis showed the utility of achene features for taxonomic groupings of the species within the studied genera. Although the specimen samples represented a limited range of taxa, the achene features and pericarp anatomy provided a reasonable source for the taxonomic treatment of the studied genera within the subtribe.
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Authors and Affiliations

Balkrishna Ghimire
1
ORCID: ORCID
Dabin Yum
2
Jae Hyeun Kim
2
Mi Jin Jeong
2

  1. Division of Forest Biodiversity, Korea National Arboretum, Pocheon 11186, Korea
  2. Division of Plant Resources, Korea National Arboretum, Pocheon 11186, Korea
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Abstract

This paper reports on 29 species of lichenicolous fungi collected in the Hornsund region and Sørkapp Land area, Spitsbergen. New to science are Hystrix gen. nov., Slellifraga gen. nov., Dactylospora cladoniicola sp. nov., Hystrix peltigericola sp. nov., Stellifraga cladoniicola sp. nov. and Zwackhiomyces macrosporus sp. nov. A further 15 species are new to Svalbard.

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Authors and Affiliations

Vagn Alstrup
Maria Olech
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Abstract

This paper reports on eleven species of hepatics collected on King George Island, South Shetland Islands (6Г50'—62°15'S latitude and 57°30'—59 00'W longitude). A short account of the vegetation of this Antarctic island is provided and the role of liverworts in particular plant communities is discussed. Two species, Hygrolembidium ventrosum (Mitt.) Grolle and Scapania abcordata (Berggr.) S. Arnell are reported for the first time from the Antarctic botanical zone; the latter is recorded for the first time in the Southern Hemisphere and, additionally, this is the first record of the genus Scapania from Antarctica. A detailed description of the habitat of each taxon is given and distribution maps for the eleven species are provided. A key to the eleven species from King George Island is given, and a detailed taxonomic discussion is included for Cephaloziella varians (Gott.) Steph and Lophozia excisa (Dicks.) Dumort. The former is considered to be synonymous with the widespread Arctic species C. arctica Bryhn & Douin ex K. Müll.

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Authors and Affiliations

Ryszard Ochyra
Jiři Váňa
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Abstract

On the basis of comparable habit, leaf morphology and leaf cell pattern, leaf and stem sectional anatomy, Dichelyma antarcticum C. Muell. is reduced to synonymy with Blindia magellanica C. Muell.

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Authors and Affiliations

Ryszard Ochyra
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Abstract

The shallow-marine carbonate deposits of the Reuchenette Formation (Kimmeridgian, Upper Jurassic) in

northwestern Switzerland and adjacent France yield highly diverse bivalve associations, but only rarely contain

remains of pinnid bivalves. The three occurring taxa Pinna (Cyrtopinna) socialis d’Orbigny, 1850, Stegoconcha

granulata (J. Sowerby, 1822) and Stegoconcha obliquata (Deshayes, 1839) have been revised. A lectotype for

Pinna (C.) socialis was designated and the taxon is assigned herein to P. (Cyrtopinna) Mörch, 1853, the first record

of the subgenus from the Jurassic. A brief review of Stegoconcha Böhm, 1907 revealed two species groups

within the genus. Species close to the type species S. granulata are characterized by a nearly smooth anterior

shell, followed posteriorly by deep radial furrows and rows of pustules covering the dorsal flank. Another group

comprises radially ribbed species related to S. neptuni (Goldfuss, 1837). It includes among others the Paleogene

species S. faxensis (Ravn, 1902), extending the known range of Stegoconcha from the Middle Jurassic into the

Paleogene. The paper suggests a relationship between Stegoconcha and the Cretaceous Plesiopinna Amano,

1956, with S. obliquata as a possible intermediate species leading to Plesiopinna during the Early Cretaceous.

Furthermore, a possible relationship between Stegoconcha and Atrina Gray, 1842 is discussed.

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Authors and Affiliations

Jens Koppka
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Abstract

Sparse fish microremains have been found in marine limestones from the Middle Devonian (Givetian) Skały

Formation (Sitka Coral-Crinoid Limestone Member and Sierżawy Member), Świętomarz–Śniadka section,

Bodzentyn Syncline, Łysogóry Region, northern Holy Cross Mountains, associated with conodonts of the

hemiansatus to ansatus zones. Thelodont scales referred here to Australolepis sp. cf. A. seddoni come from near

Śniadka village, from samples dated as hemiansatus to rhenanus/varcus zones. This increases the known range

for the genus from its original find in Western Australia. The presence of a thelodont in the late Middle Devonian

in Poland extends the known distribution of turiniids around the peri-Gondwana shorelines of Palaeotethys.

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Authors and Affiliations

Susan Turner
Michał Ginter
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Abstract

The decapod fauna from the Badenian (middle Miocene) deposits of western Ukraine comprises in total 31 taxa: 20 species, 9 taxa left in open nomenclature, and 2 determined at family level. Thirteen of these taxa are reported for the first time from the territory of Ukraine. Among them are the first records of Trapezia glaessneri Müller, 1976 in the Fore-Carpathian Basin and Pachycheles sp. in Paratethys. One taxon (Petrolisthes sp. A) probably represents a new species. The occurrence of this significant decapod fauna is restricted almost exclusively to the Upper Badenian (i.e., early Serravallian) coralgal reefs of the Ternopil Beds. The taxonomic composition of the decapods indicates that the Late Badenian depositional environment was a shallow marine basin dominated by reefs that developed in warm-to-tropical waters of oceanic salinity. The decapod assemblage from the Ternopil Beds is similar in its taxonomic composition to numerous decapod faunules from fossil reefs of Eocene to Miocene age from the Mediterranean realm and of Miocene age from Paratethys. In contrast, decapod remains are very scarce in Badenian siliciclastic deposits (Mikolaiv Beds) and are represented by the most resistant skeletal elements, i.e., dactyli and fixed fingers. This scarcity was caused by the high-energy environment, with frequent episodes of redeposition, which disintegrated and abraded the decapod remains.

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Authors and Affiliations

Marcin Górka
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Abstract

Until recently, Festuca arietina was practically an unknown species in the flora of Eastern Europe. Such a situation can be treated as a consequence of insufficient studying of Festuca valesiaca group species in Eastern Europe and misinterpretation of the volume of some taxa. As a result of a complex study of F arietina populations from the territory of Ukraine (including the material from locus classicus), Belarus and Lithuania, original anatomy, morphology and molecular data were obtained. These data confirmed the taxonomical status of F arietina as a separate species. Eleven morphological and 12 anatomical characters, ITS1-5.8S-ITS2 cluster of nuclear ribosomal genes, as well as the models of secondary structure of ITS1 and ITS2 transcripts were studied in this approach. It was found for the first time that F arietina is hexaploid (6x = 42), which is distinguished from all the other narrow-leaved fescues by specific leaf anatomy as well as in ITS1-5.8S-ITS2 sequences. Molecular data indicating possible hybridogenous origin of F arietina, fall in line with the anatomical-morphological data and explain the tendency toward sclerenchyma strands fusion with formation of a continuous ring in F arietina, as well as E arietina ecological confinement to psammophyte biotopes.

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Authors and Affiliations

I Bednarska
I Kostikov
A Tarieiev
V Stukonis
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Abstract

The 2030 Agenda for Sustainable Development was adopted in 2015. The United Nations framework does not directly include raw materials in Sustainable Development Goals (SDGs). The mining industry has a great impact not only for the environment but also for its stakeholders, both from the nearest and the most remote surrounding. As such, the mining industry has the opportunity and potential to both positively and negatively impact on all seventeen SDGs. The introduction of the EU directive on the disclosure of non-financial data has a great impact on the reporting of sustainable development reporting. Additionally, in March 2020, the European Commission published the EU Taxonomy. With regard to the current geopolitical situation, some European Union members, such as Germany, France and the Netherlands, have taken the decision to open or re-open of their coal-fired plants. Admittedly, these countries underline that the inclusion of coal in their power industry is only temporary and limited to a well-defined period of time. The implementation of the SDGs should be partially important in the case of mining, the activities of which involve the extraction of various types of mineral resources, especially non-renewable resources. This raises two fundamental research questions; what is the actual level of the reporting of SDGs in the polish mining industry, and if the EU Taxonomy Regulations will increase the reporting of SDGs in Polish mining?
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Authors and Affiliations

Olga Julita Janikowska
1
ORCID: ORCID

  1. Mineral and Energy Economy Research Institute, Polish Academy of Sciences, Kraków, Poland
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Abstract

This study investigated the potential of three microalgae taxonomic groups of Chlorophyta, Cyanoprokaryota and Bacillariophyceae for biogas production. Biogas potential was assessed in mesophilic anaerobic digestion batch tests over a period of 20 days. The cumulative biogas yield (CBY) of Chlorophyta and Cyanoprocaryota was respectively 396.21 mL/g Volatile Solids (VS) and 382.45 mL/g VS. Bacillariophyceae digestion showed lower biogas production of 357.07 mL/g VS. The highest cumulative methane yield (CMY) of 241.25 mL CH 4/g VS was recorded for Cyanoprocaryota biomass, which was signifi cantly higher (p<0.05) than the other two types of microalgae. The highest methane content in biogas of 63.08% was observed with Cyanoprokaryota. Chemical composition of biomass as well as biogas productivity are infl uenced by algal taxonomy.
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Authors and Affiliations

Marta Kisielewska
1
Marcin Dębowski
1
Marcin Zieliński
1

  1. Uniwersytet Warmińsko-Mazurski w Olsztynie, Department of Environmental Engineering
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Abstract

The known occurrence of corals distinguished here in the new Family Krynkaphyllidae varies at the subfamily level. Those of the Subfamily Krynkaphyllinae subfam. nov. are so far almost unknown from outside of the Donets Basin. In contrast, those of the Subfamily Colligophyllinae subfam. nov. are common, possibly ranging from the lower Viséan Dorlodotia Salée, 1920, a potential ancestor of the family, to the Artinskian Lytvophyllum tschernovi Soshkina, 1925. They bear different generic names, but were all originally described as fasciculate colonial. A detailed study of Lytvophyllum dobroljubovae Vassilyuk, 1960, the type species of Colligophyllum gen. nov., challenges that recognition in that at least some of those taxa are solitary and gregarious and/or protocolonial. As such, solitary, protocolonial and, probably, fasciculate colonial habits are accepted in the Colligophyllinae subfam. nov., whereas the Krynkaphyllinae subfam. nov. contains only solitary taxa. The resemblance to the Suborder Lonsdaleiina Spasskiy, 1974 led to the analysis of families included in that suborder by Hill (1981) in the context of their relationship, or homeomorphy, to Krynkaphyllidae fam. nov. This question primarily concerns the Family Petalaxidae Fomichev, 1953; a relationship with the Family Geyerophyllidae Minato, 1955, is more distant, if one exists. The distinct, parallel stratigraphic successions of taxa within two subfamilies of the Krynkaphyllidae fam. nov. document their probably common roots and early divergence. However, a lack of robust data precludes an interpretation or treatment of those successions as phylogenetic. The absence of key stratigraphic and morphologic data meant that eastern Asiatic taxa have not been considered in these successions; however, morphological similarities allow for their tentative inclusion within the Krynkaphyllidae fam. nov. The following new taxa are introduced: Krynkaphyllidae fam. nov., Krynkaphyllinae subfam. nov., Colligophyllinae subfam. nov., Krynkaphyllum gen. nov., Colligophyllum gen. nov., Protokionophyllum feninoense sp. nov., Krynkaphyllum multiplexum sp. nov., Krynkaphyllum validum sp. nov., and three species of Protokionophyllum Vassilyuk in Aizenverg et al., 1983 left in open nomenclature.
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Authors and Affiliations

Jerzy Fedorowski
1

  1. Institute of Geology, Adam Mickiewicz University, Bogumiła Krygowskiego 12, PL-61-680 Poznań, Poland
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Abstract

The Family Kumpanophyllidae Fomichev, 1953, synonymised by Hill (1981) with the Family Aulophyllidae Dybowski, 1873, is emended and accepted as valid. The new concept of this family, based on both new collections and discussion on literature data, confirms the solitary growth form of its type genus Kumpanophyllum Fomichev, 1953. However, several fasciculate colonial taxa, so far assigned to various families, may belong to this family as well. The emended genus Kumpanophyllum forms a widely distributed taxon, present in Eastern and Western Europe and in Asia. Its Serpukhovian and Bashkirian occurrences in China vs Bashkirian occurrences in the Donets Basin and in Spain, may suggest its far-Asiatic origin, but none of the existing taxa can be suggested as ancestral for that genus. Thus, the suborder position of the Kumpanophyllidae remains unknown. Four new species: K. columellatum, K. decessum, K. levis, and K. praecox, three Kumpanophyllum species left in open nomenclature and one offsetting specimen, questionably assigned to the genus, are described.

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Authors and Affiliations

Jerzy Fedorowski
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Abstract

A relatively rich assemblage of starfish is recognised from the talus facies of an Upper Jurassic (lower Kimmeridgian) biohermal, sponge-cyanobacterial build-up from the Wapienno/Bielawy succession exposed in a salt-dome anticline in Kuyavia region, north-central Poland. The paper presents 8 taxa belonging to 4 genera (one new to science): Boxaster gen. nov., Noviaster Valette, 1929, Tylasteria Valette, 1929, Valettaster Lambert, 1914, and 4 families: Astropectinidae Gray, 1840, Goniasteridae Forbes, 1841, Sphaerasteridae Schöndorf, 1906 and Stauranderasteridae Spencer, 1913. Only a very few representatives of some of these taxa have formerly been reported from the Jurassic of Poland. Two species are new: Valettaster planus sp. nov. and Boxaster wapienensis gen. et sp. nov. The Jurassic starfish assemblage recognised from the Wapienno/Bielawy succession is interpreted as an offshore starfish fauna with the admixture of allochtonous shallow-water taxa. The lithology of the source deposits indicates their transport by storm agitation and/or mass movements. This fact strongly influenced the preservation state, all collected plates being disarticulated and most of them abraded.
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Authors and Affiliations

Michał Loba
1
Urszula Radwańska
2

  1. Polish Academy of Sciences Museum of the Earth in Warsaw, Aleja na Skarpie 20/26, 27, 00-488 Warszawa
  2. Faculty of Geology, University of Warsaw, Żwirki i Wigury 93, 02-089 Warszawa, Poland
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Abstract

The paper focuses on the taxonomic description of the lower Carboniferous (uppermost Tournaisian to middle Viséan) solitary rugose corals from bedded limestone and shale units in the Flett Formation in the Jackfish Gap (eastern Liard Range), northwestern Canada. The corals described herein include 12 species representing the genera Ankhelasma Sando, 1961, Bradyphyllum Grabau, 1928, Caninophyllum Lewis, 1929, Cyathaxonia Michelin, 1847, Ekvasophyllum Parks, 1951, Enniskillenia Kabakovich in Soshkina et al., 1962, Vesiculophyllum Easton, 1944 and Zaphrentites Hudson, 1941. Two of these species are new (Ankhelasma canadense sp. nov. and Ekvasophyllum variabilis sp. nov.) and 6 taxa are described in open nomenclature. The distribution and relative abundance of solitary Rugosa in Europe and the Liard Basin confirm the geographical proximity of those areas and the open marine communication between them during the early Carboniferous. It therefore represents an important contribution to the determination of the time of isolation of the western Laurussia shelf fauna from that of southeastern Laurussia, as well as the time of the possible emergence of species from southeastern Laurussia into the western Laurussia seas. Of particular importance here are cosmopolitan taxa and the timing of their disappearance from the fossil record.
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Authors and Affiliations

Edward Chwieduk
1

  1. Institute of Geology, Adam Mickiewicz University, Bogumiła Krygowskiego 12, 61-680 Poznań, Poland
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Abstract

The common and ecologically important cyanobacterial form-genus Leptolyngbya is widely distributed in numerous ecosystems over the Earth's biosphere. Several morphospecies dominate microbial communities in polar habitats, but their diversity and local ecological significance are little known. Several articles characterising strains isolated from Antarctic coastal habitats by molecular methods were published, but knowledge of their phenotype and ecological characters are indispensable for future detailed environmental studies. Distinct morpho- and ecotypes (ecologically important morphospecies) from maritime Antarctica are characterised in this article. Eight dominant Leptolyngbya types from subaerophytic and freshwater habitats were recognised, and four of them (L. borchgrevinkii, L. fritschiana, L. nigrescens and L. vincentii) are described as new distinct species.

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Authors and Affiliations

Jiří Komárek
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Abstract

Grimmia andreaeopsis C. Muell., a species described from sterile material from the Chukotka Peninsula, is redescribed and illustrated The species is actually a member of the genus Schistidium. It can be distinguished from its closest relatives, viz. species of S. strictum complex, by the possession of a unique combination of characters: (1) inky black coloration of gametophytes; (2) strongly and asymmetrically keeled, rapidly wide-spreading to squarrose when moist, leaves; (3) cells entirely smooth, very incrassate and strongly nodulose nearly to the base of the lamina: (4) a costa totally smooth or only occasionally slightly roughened on the back below the apex, but never scabrous with conical papillae; (5) leaf margins always entire; (6) peristome teeth bluntly acuminate. Unlike most rupestral species of Schistidium it grows in wet arctic fens. S. holmenianum Steere & Brassard, a species known to be widely distributed in the Nearctic, and Racomitrium depressum Lesq. var. nigricans Kindb., a variety described from Labrador and Hudson Bay. are synonymous with S. andreaeopsis (C. Muell.) Laz. A comparison of S andreaeopsis with the Andean-Subantarctic S. anqustifolium (Mitt.) Herz is made and these species are considered to be closely related, but not conspecific, bipolar counterparts. Also, a comparison with the South Georgian S. urnulaceum (C. Muell.) Bell and the Holarctic species of S. strictum complex, which are characterized by having similar leaf cell patterns, is made. S. andreaeopsis has a circumpolar distribution, mainly within the High Arctic. In addition to the Nearctic, the species is known to occur in Svalbard, North Land, Taymyr Peninsula, Yakutia, Wrangel Island, and on the Chukotka Peninsula.

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Authors and Affiliations

Ryszard Ochyra
Olga M. Afonina

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