Trace fossils Lockeia siliquaria James, Ophiomorpha nodosa Lundgren, Parataenidium seymourensis isp. n., Protovirgularia rugosa (Miller and Dyer) and Rhizocorallium jenense Zenker have been described for the first time from the Eocene La Meseta Formation of Seymour (Marambio) Island, Antarctic Peninsula . Determinations of some trace fossils formerly described from this formation have been revised. The whole trace fossils association from the La Meseta Formation points to foreshore-offshore environment as indicated by presence of the Skolithos and Cruziana ichnofacies, and to at least temporal normal salinity.
Body size is an important measure in biology and especially in paleobiology. With respect to fossil penguins from the Eocene La Meseta Formation of Seymour Island (West Antarctica) the overall size has to be judged from the dimensions of single bones. The analysis based on selected measurements of hind limb bones from the Polish collection of Eocene Antarctic penguins yielded results supporting predictions published formerly. Estimated body masses and lengths indicate that mean interspecific body size of extinct Antarctic Spheniscidae exceeded that of Recent species.
One of the most significant global climatic events in the Cenozoic was the transition from greenhouse to icehouse conditions in Antarctica. Tectonic evolution of the region and gradual cooling at the end of Eocene led to the first appearance of ice sheets at the Eocene/Oligocene boundary (ca. 34 Ma). Here we report geological record of mountain glaciers that preceded major ice sheet formation in Antarctica. A terrestrial, valley-type tillite up to 65 metres thick was revealed between two basaltic lava sequences in the Eocene– Oligocene Point Thomas Formation at Hervé Cove – Breccia Crag in Admiralty Bay, King George Island, South Shetland Islands. K-Ar dating of the lavas suggests the age of the glaciation at 45–41 Ma (Middle Eocene). It is the oldest Cenozoic record of alpine glaciers in West Antarctica, providing insight into the onset of glaciation of the Antarctic Peninsula and South Shetland Islands.
Early Palaeocene through early Eocene silicoflagellate assemblages were examined from five southern subtropical
through subpolar deep-sea sites: DSDP Holes 208 and 524, and ODP Holes 700B, 752A, and 1121B. For each
site, the taxonomic composition of the silicoflagellate assemblage is documented in detail; Pseudonaviculopsis
gen. nov., Dictyocha castellum sp. nov. and Stephanocha? fulbrightii sp. nov. are proposed, along with several
new combinations. More importantly, however, these observations enable a considerable refinement to the existing
Palaeocene–Eocene silicoflagellate biostratigraphic zonation that for the first time uses datums calibrated to
the Geomagnetic Polarity Timescale. The Corbisema aspera Interval Zone occurs immediately above the K/Pg
boundary and is here described from Seymour Island. The Corbisema hastata Partial Range Zone extends from
near the K/Pg boundary to late early Palaeocene and has been observed in Hole 208. The Pseudonaviculopsis disymmetrica
Acme Zone occurs in Holes 208 and 700B. The Dictyocha precarentis Partial Range Zone, observed
in Holes 208, 700B, 752A and 1121B, is subdivided into D. precarentis, Naviculopsis primativa, N. cruciata
and Pseudonaviculopsis constricta subzones. The Naviculopsis constricta Partial Range Zone occurs in Holes
524, 700B, 752A and 1121B. This study is also the first to consider syn- and/or diachroneity in Palaeogene
silicoflagellate biostratigraphy.
A few specimens of a macroporid bryozoan were collected, from the Eocene La Meseta Formation from Seymour (Marambio) Island, Antarctic Peninsula. Based on the morphology of the studied specimens Macropora antarctica sp.n. has been erected. This is the stratigraphically oldest species of the genus which exhibits a number of similarities with the Tertiary fossils and some Recent macroporids reported from the Southern Hemisphere i.e., Australia, New Zealand and South America.
Eocene penguin remains from Seymour Island (Antarctica) are so far the oldest−known record of extinct Sphenisciformes. Rich Argentineand Polish collections of penguin bones from the La Meseta Formation are taxonomically revised on tarsometatarsal morphology. Two genera and four species are erected: Mesetaornis polarisgen. et sp. n., Marambiornis exilisgen. et sp. n., Delphinornis arctowskiisp. n. and D. gracilissp. n. Moreover, the diagnoses of already described species: Anthropornis nordenskjoeldi, A. grandis, Palaeeudyptes klekowskii, P. gunnari, Archaeospheniscus wimani and Delphinornis larseniare revised as well. Gradual cooling of climate, changes of environment andtrophic relationships, that lasted several millions years, were most probably responsible forthe intense speciation and taxonomic diversification of the Middle–Late Eocene La Meseta penguins.
Skeletal remains of penguins from the Eocene La Meseta Formation (Seymour Island, Antarctica) constitute the only extensive fossil record of Antarctic Sphenisciformes. No articulated skeletons are known, and almost all fossils occur as single isolated elements. Most of the named species are based on tarsometatarsi (for which the taxonomy was revised in 2002). Here, 694 bones (from the Polish collection) other than tarsometatarsi are reviewed, and allocated to species. They confirm previous conclusions and suggest that ten species grouped in six genera are a minimal reliable estimate of the Eocene Antarctic penguin diversity. The species are: Anthropornis grandis, A. nordenskjoeldi, Archaeospheniscus wimani, Delphinornis arctowskii, D. gracilis, D. larseni, Marambiornis exilis, Mesetaornis polaris, Palaeeudyptes gunnari and P. klekowskii. Moreover, diagnoses of four genera (Anthropornis, Archaeospheniscus, Delphinornis and Palaeeudyptes) and two species (P. gunnari and P. klekowskii) are supplemented with additional, non-tarsometatarsal features. Four species of the smallest penguins from the La Meseta Formation (D. arctowskii, D. gracilis, M. exilis and M. polaris) seem to be the youngest taxa within the studied assemblage - their remains come exclusively from the uppermost unit of the formation. All ten recognized species may have co-existed in the Antarctic Peninsula region during the Late Eocene epoch.
Penguin bones from the La Meseta Formation (Seymour Island, Antarctic Peninsula) are the only record of Eocene Antarctic Sphenisciformes. Being an abundant component of the youngest unit of the formation (Telm7), they are not so common in earlier strata. Here, I present the oldest penguin remains from the La Meseta Formation (Telm1-Telm2), often bearing close resemblance to their counterparts from younger units. Addressing the recent findings in fossil penguin systematics, I suggest there is too weak a basis for erecting new Eocene Antarctic taxa based on non-tarsometatarsal elements of penguin skeletons, and considering Oligocene species part of the studied assemblage. Finally, I conclude if the common ancestor of extant Sphenisciformes lived in the Eocene Antarctic (as suggested recently), penguins referred to Delphinornis seem to be prime candidates to that position.
Isolated and fragmented jaws, a single basioccipitale and vertebrae of the Gadiformes, indeterminate family and genus, are described from Eocene sediments of the La Meseta Formation, Seymour Island, Antarctic Peninsula. Based on the dentition and other characters of both jaws they are assigned an informal name of „Mesetaichthys". The remaining isolated bones belong probably to the same form.
Pyrite framboids occur in loose blocks of plant−bearing clastic rocks related to volcano−sedimentary succession of the Mount Wawel Formation (Eocene) in the Dragon and Wanda glaciers area at Admiralty Bay, King George Island, West Antarctica. They were investigated by means of optical and scanning electron microscopy, energy−dispersive spectroscopy, X−ray diffraction, and isotopic analysis of pyritic sulphur. The results suggest that the pyrite formed as a result of production of hydrogen sulphide by sulphate reducing bacteria in near surface sedimentary environments. Strongly negative δ34SVCDT values of pyrite (−30 – −25 ‰) support its bacterial origin. Perfect shapes of framboids resulted from their growth in the open pore space of clastic sediments. The abundance of framboids at cer− tain sedimentary levels and the lack or negligible content of euhedral pyrite suggest pulses of high supersaturation with respect to iron monosulphides. The dominance of framboids of small sizes (8–16 μm) and their homogeneous distribution at these levels point to recurrent development of a laterally continuous anoxic sulphidic zone below the sediment surface. Sedimentary environments of the Mount Wawel Formation developed on islands of the young magmatic arc in the northern Antarctic Peninsula region. They embraced stagnant and flowing water masses and swamps located in valleys, depressions, and coastal areas that were covered by dense vegetation. Extensive deposition and diagenesis of plant detritus in these environments promoted anoxic conditions in the sediments, and a supply of marine and/or volcanogenic sulphate enabled its bacterial reduction, precipitation of iron mono− sulphides, and their transformation to pyrite framboids.
Until now Eocene chimeroid holocephalians of Antarctica have been known from only a few specimens attributed to two species from the Eocene of Seymour Island. New material collected by Polish and English field parties includes numerous tooth plates and fin spine fragments from the Eocene La Meseta Formation. We describe a new species, Callorhinchus stahli, based on two mandibular and a single fragmentary palatine tooth plate. In addition, the stratigraphic distribution and diversity of Eocene Antarctic chimeroids is discussed. The chimeroid Ischyodus shows the greatest stratigraphic distribution with its greatest abundance in the middle parts of the La Meseta Formation while Chimaera and Callorhinchus are restricted to the lower ones. Changes in the environment and habitat availability most probably triggered the distributional pattern and the disappearance of chimeroids.
An isolated, deciduous incisor of an archaic whale found in the upper part of the La Meseta Formation (Telm7) is tentatively assigned to the Archaeoceti. The strata from which the tooth was recovered are of Late Eocene (Priabonian) age, and previous reports indicate that they contain the remains of Dorudontinae (Archaeoceti) and Llanocetidae (Mysticeti). The tooth is similar in shape, size and ornamentation to the milk teeth of Zygorhiza. The enamel is mostly prismatic, with prism sheats generally open, except for the outermost layer, which is aprismatic. The Schmelzmuster consists of radial and decussating enamel types. The decussating zone has distinct Hunter-Schreger bands (HSB), usually consisting of 1012 prisms. It is bordered by an external zone built of radial enamel extending for 22% of the enamel thickness and an internal, starting zone, with less developed HSB, occupying 9% of the enamel thickness. The interprismatic matrix is parallel to the prism direction. An archaeocete origin of the tooth is suggested by its enamel features, typical for the group. However, additional study of the Llanocetidae enamel structure is needed for final identification.